UWB Crest

The Thorpe Group



Creer, S., Pook, C.E., Malhotra, A., Thorpe, R.S. 2006. Optimal intron analyses in the Trimeresurus radiation of Asian pitvipers. Systematic Biology 55: 57-72. PDF

Gow JL, H Johansson, Y Surget-Groba, R S Thorpe. 2006. Ten polymorphic tetranucleotide microsatellite markers isolated from the Anolis roquet series of Caribbean lizards. Molecular Ecology Notes. 6:873-876.  PDF

Sanders KL,  A. Malhotra, R.S. Thorpe.  2006. Evidence for a Müllerian mimetic radiation in Asian pitvipers. Proceedings of the Royal Society B 273:1135-1141. PDF

Sanders KL,  A. Malhotra, R.S. Thorpe.  2006. Combining molecular,morphological and ecological data to infer species boundaries in a cryptic tropical pitviper.Biological Journal of the Linnean Society. 87: 343-664. PDF


Thorpe RS. 2005. Population evolution and island biogeography. Science. 310:1778-1779. PDF
Science summary link http://www.sciencemag.org/cgi/content/summary/310/5755/1778?ijkey=jJvlLKRj4UDc6&keytype=ref&siteid=sci
Science Full text link  http://www.sciencemag.org/cgi/content/full/310/5755/1778?ijkey=jJvlLKRj4UDc6&keytype=ref&siteid=sci

Creer S, A Malhotra, R S Thorpe, C E Pook. 2005. Targetting introns for phylogenetic analyses in non-model taxa: experimental results in Asian pitvipers. Cladistics. 21:390-395. PDF

Thorpe, RS  DL Leadbeater, C E Pook. 2005. Molecular clocks and geological dates: cytochrome b of Anolis extremus substantially contradicts dating of Barbados emergence. Mol Ecol 14:2087-2096. PDF

Thorpe RS, JT Reardon, A Malhotra. 2005. Common garden and natural selection experiments support ecotypic differentiation in the Dominican anole (Anolis oculatus). Amer Nat . 165:495-504. PDF

Wüster W, JE Ferguson, JA Quijada-Mascareñas, CE Pook, MG Salomão & RS Thorpe 2005 Tracing an invasion: landbridges, refugia and the phylogeography of the Neotropical rattlesnake (Serpentes: Viperidae: Crotalus durissus) .  Mol Ecol. 14: 1095-1108. PDF

Gübitz T, RS Thorpe, A Malhotra  2005. The dynamics of genetic and morphological variation on volcanic islands Proc R. Soc B. 272:751-757. PDF

Malhotra A, & RS Thorpe. 2005. Erratum to “A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis)” {Mol Phylogenet. Evol. 32 (2004) 83-100]. Mol Phylo. Evol. 34:68-681. PDF


Stenson,AG RS Thorpe and A Malhotra. 2004. Evolutionary differentiation of bimaculatus group anoles based on analyses of mtDNA and microsatellite data. Mol Phylo Evol 32:1-10. PDF

Malhotra, A., and R.S.Thorpe. 2004. A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pit vipers (Trimeresurus and Ovophis). Mol Phylo Evol. 32: 83-100. PDF

Creer, S., Thorpe, R.S., Malhotra, A.,  W.-H. Chou. & AG Stenson 2004. The utility of AFLPs for supporting mitochondrial DNA phylogeographical analysres in the Taiwanese bamboo viper, Trimeresurus stejnegeri. J. Evol. Biol. 17: 100-107. PDF

Sanders, K.L., A. Malhotra, and R.S. Thorpe. 2004. Ecological diversification in a group of Indomalayan pitvipers (Trimeresurus): convergence in taxonomically important traits has implications for species identification. J. Evol. Biol. 17: 721-731. PDF

Herrmann, H.W., T. Ziegler, A. Malhotra, R.S. Thorpe, and C.L. Parkinson. 2004. Redescription and systematics of Trimeresurus cornutus (Serpentes: Viperidae) based on morphology and molecular data. Herpetologica, 60: 211-221. PDF

Malhotra, A., and R.S. Thorpe. 2004. Maximising information in systematic revisions: a combined molecular and morphological analysis of a cryptic green pitviper complex (Trimeresurus stejnegeri). Biol. J. Linn. Soc., 82: 219-235. PDF

Malhotra, A., R.S. Thorpe and B.L. Stuart. 2004. A morphometric analysis of Trimeresurus vogeli (David, Vidal and Pauwels, 2001), with new data on diagnostic characteristics, distribution and natural history. Herpetological Journal. 14: 65-77. PDF

Malhotra, A., and R.S. Thorpe. 2004. Reassessment of the validity and diagnosis of the pitviper Trimeresurus venustus Vogel, 1991 Herpetological Journal, 14: 21-33. PDF

Sanders, K.L., A. Malhotra, A. Gumprecht, R. S. Thorpe & U. Kuch. 2004. Poeia inornata, a new species of pitviper from West Malaysia (Squamata: Viperidae: Crotalinae). Russian J. Herpetol.11: 171-184. PDF

Thorpe RS, Malhotra A, Stenson AG and Reardon JT (2004)  Adaptation and speciation in Lesser Antillean anoles. In: (eds.) Dieckmann, U., Doebeli, M., Metz JAJ,  and Tautz, D., (Eds.) pp322-344  In:  Adaptive Speciation,  Cambridge University Press, U.K. PDF


Thorpe RS. 2003. Garth Underwood (1919-2002): A vision of reptile systematics. Herpetological Review. 34:6-8. PDF

Thorpe RS and Stenson AG (2003)  Phylogeny, paraphyly and ecological adaptation of the colour and pattern in the Anolis roquet complex on Martinique.  Mol. Ecol.  12, 117-132. PDF

Creer S, Malhotra A and Thorpe RS (2003)  Assessing the phylogenetic utility of four mitochondrial genes and a nuclear intron in the Asian pitviper genus Trimeresurus:  separate, simultaneous, and conditional data combination analyses.  Mol Biol Evol 20, 1240-1251. PDF

Creer S, Malhotra A, Thorpe RS, Stocklin R, Favreau P and Chou W-H (2003)  Genetic and ecological correlates of intraspecific variation in pitviper venom composition detected using matrix assisted laser desorption time-of-flight mass spectrometry (MALDI-TOF-MS) and isoelectric focusing.  J Mol Evol  56, 317-329. PDF


Ogden R, Thorpe RS (2002). Molecular evidence for ecological speciation in tropical habitats. PNAS 99: 13612-13615.  PDF

Thorpe RS (2002). Analysis of Color Spectra in Comparative Evolutionary Studies: Molecular Phylogeny and Habitat Adaptation in the St. Vincent Anole (Anolis trinitatis). Syst Biol 51: 554-569.  PDF

Ogden R, Thorpe RS (2002). The usefulnes of amplified fragment length polymorphism markers for taxon discrimation across graduated fine evolutionary levels in Caribbean Anolis lizards. Mol. Ecol. 11: 437-445. PDF

Stenson AG, Malhotra A, Thorpe RS (2002). Population differentiation and nuclear gene flow in the Dominican anole (Anolis oculatus). Mol. Ecol. 11: 1679-1688. PDF

Creer, S., Chou, W.-H., Malhotra, A. and Thorpe, R.S. 2002. Offshore insular variation in the diet of the Taiwanese bamboo viper, Trimeresurus stejnegeri (Schmidt). Zoological Science, 19:907-913. PDF

Sanders, K.L., Malhotra, A., and R.S. Thorpe. 2002. A contribution to the systematics of two commonly confused pitvipers from the Sunda region: Trimeresurus hageni and T. sumatranus. Bull. Nat. Hist. Mus, Zoology, 68(2):107-111. PDF

Hille A, Janssen IAW, Menken SBJ, Schlegel M, Thorpe RS (2002). Heterologous Amplification of Microsatellite Markers From Colubroid Snakes in European Natricines (Serpentes : Natricinae). Journal of Heredity 93: 63-66.  PDF

Ogden R, Griffiths TJ, Thorpe RS (2002). Eight Microsatellite Loci in the Caribbean Lizard, Anolis roquet. Conservation Genetics 3: 345-346. PDF

Thorpe RS (2002)  Geographic variation. In: Pagel MD,  (Ed.)  Encyclopedia of Evolution, pp. 430-435. U.K.Oxford University Press PDF

Wuster W, Salomao MG, Quijada-Mascarenas JA, Thorpe RS. 2002. Origins and evolution of the South American pit viper fauna: evidence from mitiochondrial DNA sequence analysis. In. Biology of the Vipers (eds Schuett GW, Hoggren M, Douglas ME, Greene HW) pp 111-128 Eagle Mountain Publishing, Eagle Mt, Utah. PDF

Wüster W, Thorpe RS Salomao MG, Thomas, L., Puorto G, Theakston RDG, & Warrell, DA. 2002. Origin and phylogenetic position of the Lesser Antillean species of Bothrops (Serpentes, Viperidae): biogeographical and medical implications. Bull. Nat. Hist. Mus, Zoology, 68: 101-106. PDF


Thorpe RS, Richard M (2001). Evidence That Ultraviolet Markings Are Associated With Patterns of Molecular Gene Flow. PNAS 98: 3929-3934. PDF

Creer S, Malhotra A, Thorpe RS, Chou WH (2001). Multiple Causation of Phylogeographical  Pattern as Revealed by Nested Clade Analysis of the Bamboo Viper (Trimeresurus stejnegeri) Within Taiwan. Mol. Ecol. 10: 1967-1981. PDF

Giannasi N, Malhotra A, Thorpe RS (2001). Nuclear and MtDNA Phylogenies of the Trimeresurus Complex: Implications for the Gene Versus Species Tree Debate. Mol Phylo Evol 19: 57-66. PDF

Giannasi N, Thorpe RS, Malhotra A (2001). The Use of Amplified Fragment Length Polymorphism in Determining Species Trees at Fine Taxonomic Levels: Analysis of a Medically Important Snake, Trimeresurus albolabris. Mol. Ecol.  10: 419-426. PDF

Richard M, Thorpe RS (2001). Can microsatellites be used to infer phylogenies? Evidence from population affinities of the western Canary Island lizard (Gallotia galloti).  Mol Phylo Evol 20: 351-360. PDF

Surget-Groba Y, Heulin B, Guillaume G P, Thorpe RS, Kupriyanova L, Vogrin N, Maslak R, Mazzotti S, Venczel M, Ghira I, Odierna G, Leontyeva O, Monney JC, Smith N (2001). Intraspecific Phylogeography of Lacerta vivipara and the Evolution of Viviparity. Mol Phylo Evol 18: 449-459. PDF

Puorto G, Salomao MD, Theakston RDG, Thorpe RS, Warrell DA, Wüster W (2001). Combining Mitochondrial DNA Sequences and Morphological Data to Infer Species Boundaries: Phylogeography of Lanceheaded Pitvipers in the Brazilian Atlantic Forest, and the Status of Bothrops pradoi (Squamata : Serpentes : Viperidae). J Evol Biol 14: 527-538. PDF


Richard M & Thorpe R S. 2000.Highly polymorphic microsatellites in the lacertid Gallotia galloti from the western Canary Islands. Mol Ecol.9:1919-1920. PDF

Stenson A, Malhotra A, Thorpe R S. 2000. Highly polymorphic micriosatellite loci. from the Dominican anole (Anolis oculatus) and their amplification in other bimaculatus series anoles. Mol Ecol. 9:1661-1686. PDF

Malhotra A & Thorpe R S. 2000. A phylogeny of the Trimeresurus group of pit-vipers: new evidence from a mitochondrial gene tree. Mol  Phylo Evol. 16:199-211. PDF

Giannasi, N, Thorpe R S, & Malhotra A. 2000. A phylogenetic analysis of body size evolution in the Anolis roquet group (Sauria: Iguanidae) character displacement or size assortment? Mol Ecol. 9:193-202. PDF

Gübitz T, Thorpe R S, Malhotra A, 2000. Phylogeography and natural selection in the Tenerife gecko Tarentola delalandii: testing historical and adaptive hypotheses. Mol Ecol. 9:1213-1221. PDF

Tarkhnishvili D, Thorpe R S , Arntzen JW 2000. Pre-Pleistocene refugia and differentiation between populations of the Caucasian salamander (Mertensiella caucasica). Mol Phylo Evol. 14:414-422. PDF

Pook, C E, Wüster W, Thorpe, R S. 2000. Historical biogeography of the Western Rattlesnake (Serpentes:Viperidae: Crotalus viridis) inferred from mitochondrial DNA sequence information. Mol. Phyl. Evol. 15:269-282. PDF

Malhotra A & R S Thorpe. 2000. The dynamics of natural selection and vicariance in the Dominican anole: patterns of within-island molecular and morphological divergence. Evolution 54:245-258. PDF

Brede EG, Thorpe RS, Arntzen JW, Langton TES. 2000. A morphometric study of a hybrid newt population (Triturus cristatus/T.carnifex): Beam Brook Nurseries, Surrey, U.K.  Biological Journal of the Linnean Society, 70:685-695. PDF


Arntzen, JW, Thorpe R S. 1999 Italian crested newts (Triturus carnifex) in the Basin of Geneva: distribution and genetic interactions with autochonous species. Herpetologica. 55:423-433. PDF

Malhotra, A. & R S Thorpe. 1999. Reptiles and Amphibians of the Eastern Caribbean. MacMillan  London. Info PDF

Tarkhnishvili, DN, JW Arntzen & RS Thorpe 1999 Morphological variation in brown frogs from the Caucasus and the taxonomy of the Rana macrocnemis group. Herpetologica. 55:406-417. PDF

Wüster W, Salomao MG, Duckett GJ, Thorpe RS. 1999. Mitochondrial DNA evolution of the Bothrops atrox species complex (Squamata: Serpentes: Viperidae). Kaupia 8:135-144. PDF

Salomao MG, Wüster W, Thorpe RS, 1999. MtDNA evolution in neotropical pitvipers of the genus Bothrops (Sqamata: Serpentes: Viperidae). Kaupia 8: 127-134. PDF

Wüster W, JC Daltry, & RS Thorpe. 1999. Can diet explain intraspecific venom variation? Reply to Sasa. Toxicon 37:253-258. PDF


Thorpe RS & A Malhotra. 1998. Molecular and morphological evolution within small islands. pp 67-82 in PR Grant Ed. “Evolution on islands”. Oxford University Press. PDF

Daltry JC, T Ross, RS Thorpe, W Wüster. 1998. Evidence that humidity influences snake activity patterns: a field study of the Malayan pit viper Calloselasma rhodostoma. Ecography 21:25-34. PDF

Daltry JC, Wüster, W, Thorpe RS. 1998. Intraspecific variation in feeding ecology of the crotaline snake Calloselasma rhodostoma in Southeast Asia. Journal of Herpetology 32:198-205. PDF

Saino N, Wuster W, Thorpe RS. 1998. Congruence between morphological variation and altitudinal gradient across a hybrid zone between carrion and hooded crows. Italian Journal of Zoology 65: 407-412

Raybould AF, Mogg RJ, Aldham C, Gliddon CJ, Thorpe RS, Clarke RT. 1998. The genetic structure of sea beet (Beta vulgaris ssp. maritima) populations. III. Detection of isolation by distance at microsatellite loci. Heredity 80:127-132.


Thorpe, R S, W Wüster, & A Malhotra. 1997 Venomous Snakes: Ecology, Evolution and Snakebite. The Zoological Society of London. Clarendon Press Oxford. 276pp. Info PDF

Salomao, M D G, Wüster, W., Thorpe, RS. & Touzet JM. 1997.  DNA evolution of South American pit vipers of the genus Bothrops (Reptilia: Serpentes: Viperidae). Symp. zool. Soc. Lond. 70:89-98. PDF

Malhotra, A. & Thorpe R S. 1997. New perspectives on the evolution of South-east Asian pit vipers (genus Trimeresurus) from molecular studies. Symp. zool. Soc. Lond. 70:115-128. PDF

Daltry, J, Wüster, W, & Thorpe R S. 1997. The role of ecology in determining venom variation in the Malayan pit viper, Calloselasma rhodostoma. Symp. zool. Soc. Lond. 70:155-171. PDF

Wüster, W. Salomao, MG, & Thorpe RS, Puorto G., Furtado MFD., Hoge SA, Theakston RDG., & Warrell, DA. 1997. Systematics of the Bothrops atrox complex: new insights from multivariate analysis and mitochondrial DNA sequence information. Symp. zool. Soc. Lond. 70:99-113. PDF

Belt, PJ., Malhotra A., Thorpe RS., Warrell DA, & Wüster W. 1997. Russell's viper in Indonesia: snakebite and systematics. Symp. zool. Soc. Lond. 70:219-234. PDF

Malhotra, A. & Thorpe R S. 1997. Microgeographic variation in scalation of Anolis oculatus (Dominica, West Indies): a multivariate analysis. Herpetologica. 53:49-62. PDF

Malhotra, A. & Thorpe R S. 1997. Size and shape variation in a Lesser Antillean anole, Anolis oculatus (Sauria:Iguanidae) in relation to habitat. Biol. J. Linn Soc.60:53-72. PDF

Overton, J.L., D J Macintosh & R S Thorpe. 1997. Multivariate analysis of the mud crab Scylla serrata (Brachyura:Portunidae) from four locations in Southeast Asia. Marine Biology. 128:55-62. PDF

Giannasi, N., R S Thorpe & A Malhotra. 1997. Introduction of Anolis species to the island of St Lucia, West Indies: Testing for hybrids using multivariate morphometrics. J.Herpetol. 31:586-589.  PDF


Daltry, JC, Wüster, W, & Thorpe R S. 1996. Diet and snake venom evolution. Nature. 379:537-540. PDF

Thorpe, R. S. 1996. The use of DNA divergence to help determine the correlates of evolution of morphological characters. Evolution. 50:524-531. PDF

Malhotra, A., Thorpe, R. S.,  H. Black, J. C. Daltry, and W. Wüster. 1996. Relating geographic patterns to phylogenetic processes. In "New uses for new phylogenies" ed P. H. Harvey, A. J. Leigh-Brown, J. Maynard Smith & S. Nee. Oxford University Press, Oxford. pp 187-202.

Wüster, W., R.S. Thorpe, G. Puorto. 1996. Systematics of the Bothrops atrox complex (Reptilia: Serpentes: Viperidae) in Brasil: a multivariate analysis. Herpetologica, 52:263-271. PDF

Daltry, J., G.Ponnudurai, C K Shin, N H Tan, R S Thorpe, W Wüster. 1996. Electrophoretic profiles and biological activities: Intraspecific variation in the venom of the Malayan pit viper (Callosellasma rhodostoma). Toxicon. 34:67-79. PDF

Thorpe, R S , H Black & A Malhotra. 1996. Matrix correspondence tests on the DNA phylogeny of the Tenerife Lacertid elucidates both historical causes and morphological adaptation. Systematic Biology. 45:335-343. PDF

Thorpe, R S & A Malhotra. 1996. Molecular and morphological evolution within small islands.  Philosophical Transactions of the Royal Society Lond. B. 351:815-822. PDF

Thorpe, R. S & A Malhotra. 1996. Evolutionary ecology of lizards (review). T.R.E.E. 11:523.


Thorpe, R.S., A. Malhotra, H. Black, J.C. Daltry & W. Wüster 1995. Relating geographic pattern to phylogenetic process. Philosophical Transactions of the Royal Society Lond B. 349:61-68. PDF

Thorpe, R.S., J.Daltry, W. Wüster, A. Malhotra and N.L. Thorpe. 1995. The biology of dangerous snakes: systematics and venom evolution. Toxicon 35: 589. PDF

Wüster, W., R.S. Thorpe, M.J. Cox, P. Jintakune & J. Nabhitabhata. 1995. Population systematics of the snake genus Naja (Reptilia: Serpentes: Elapidae) in Indochina: multivariate morphometrics and comparative mitochondrial DNA sequencing (cytochrome oxidase I). Journal of Evolutionary Biology 8:493-510. PDF

Malhotra , A. and R. S. Thorpe. 1995. Ameiva fuscata. Catalogue of American Amphibians and Reptiles. Ed. A Price. Society for the Study of Amphibians and Reptiles. 606:1-3. PDF


Wüster, W. and Thorpe, R. S. 1994. Naja siamensis, a cryptic species of dangerously venomous snake revealed by mtDNA sequencing. Experientia. 50:75-79. PDF

Thorpe, R.S., D.P. McGregor, A.M. Cumming and W.C. Jordan. 1994. DNA evolution and colonization sequence of island lizards in relation to geological history: mtDNA RFLP, cytochrome b, cytochrome oxidase, 12s rRNA sequence, and nuclear RAPD analysis. Evolution. 48:230-240. PDF

Thorpe, R.S., R. P. Brown, M. Day, A. Malhotra, D. P. McGregor & W. Wuster. 1994. Testing ecological and phylogenetic hypotheses in microevolutionary studies. In  Phylogenetics and EcologyEd P. Eggleton and R. Vane-Wright. Academic Press. pp 189-206. PDF

Malhotra, A. and Thorpe, R. S. 1994. Parallels between island lizards suggests selection on mitochondrial DNA and morphology. Proc Roy Soc B. 257:37-42. PDF

Castellano, S., Malhotra, A. and Thorpe, R. S. 1994. Within-island geographic variation of the dangerous Taiwanese snake Trimerusurus stejnegeri, in relation to ecology. Biol J Linn Soc. 52:365-375.  PDF

Pierce, G J., R S Thorpe, L C Hastie, A S Brierley, A Guerra, P R Boyle, R Jameison & P Avila. 1994. Geographic variation in Loligo forbesi in the Northeast Atlantic Ocean: Analysis of geographic variation and tests of causal hypotheses. Marine Biology 119:541-547. PDF

Pierce, G J., L C Hastie, A Guerra, R S Thorpe, F G Howard, P R Boyle. 1994. Morphometric variation in Loligo forbesi and Loligo vulgaris: regional, seasonal, sex, maturity and worker differences. Fisheries Research 21:127-148. PDF


Thorpe, R.S. & M. Baez. 1993. Geographic variation in scalation of the lizard Gallotia stehlini within the island of Gran Canaria. Biol. J. Linn. Soc. 48:75-87. PDF

Thorpe, R.S., McGregor, D. & Cumming, A.M. 1993. Population evolution of western Canary Island lizards (Gallotia galloti): 4‑base endonuclease restriction fragment length polymorphisms of mitochondrial DNA. Biol. J. Linn. Soc. 49:219-227. PDF

Malhotra, A. & Thorpe, R.S. 1993. An experimental field study of an eurytopic anole, Anolis oculatus. J. Zool. Lond. 229: 163-170. PDF

Thorpe, R.S., McGregor, D. & Cumming, A.M. 1993. Molecular phylogeny of the Canary Island lacertids (Gallotia): mitochondrial DNA restriction fragment divergence in relation to sequence divergence and geological time. J. Evol. Biol. 6:725-735. PDF

Brown, R. P., Thorpe, R. S. & Baez, M. 1993. Patterns and causes of morphological population differentiation in the Tenerife skink, Chalcides viridanus. Biological Journal of the Linnean Society. 50:313-328. PDF


Malhotra , A. and R. S. Thorpe. 1992. Anolis oculatus. Catalogue of American Amphibians and Reptiles. Ed. A Price. Society for the Study of Amphibians and Reptiles.540:1-4. PDF

Wüster, W. and R. S. Thorpe. 1992. Asiatic cobras: Population systematics of the Naja naja species complex (Serpentes:Elapidae) in India and Central Asia. Herpetologica. 48:69‑85. PDF

Wüster, W., S. Otsuka, R. S. Thorpe, and A. Malhotra. 1992. Morphological variation in Russell's viper in Burma and Thailand. The Herpetological Journal. 2:99‑101. PDF

Wüster, W. & R.S. Thorpe. 1992. Dentitional phenomena in Cobras revisited: spitting and fang structure in the Asiatic species of Naja  (Serpentes: Elapidae). Herpetologica: 48:424‑434. PDF

Wüster, W., S. Otsuka, A. Malhotra and R.S. Thorpe. 1992. Population systematics of Russell's viper: a multivariate study. Biol. J. Linn. Soc. 47:97‑113. PDF

Thorpe, R.S. & Malhotra, A. 1992. Are Anolis  lizards evolving? Nature, 355:506. PDF

Brown, R.P., Thorpe, R.S. & Speakman, JR. 1992. Body size, field energetics and water flux in a skink (Chalcides sexlineatus). Can. J. Zool. 70:1001-1006.

Day, M. and R S Thorpe. 1992. The Lesser Antillean Iguana, Iguana delicatissima, on St. Barthelemy. Herpetofauna News. 2:11.

Allegrucci, G., Baldari, F., Caesaroni, D., Thorpe, R.S. & Sbordoni, V. 1992. Morphometric analysis of interspecific and microgeographic variation of crayfish from a Mexican cave. Biol. J. Linn. Soc. 47:455-468.


Brown, R.P., R.S. Thorpe & M. Baez. 1991. Parallel within‑island microevolution of lizards on neighbouring islands. Nature. 352:60‑62. PDF

Malhotra, A. & R.S. Thorpe. 1991. Experimental detection of rapid evolutionary response in natural lizard populations. Nature. 353:347‑348. PDF

Thorpe, R.S. and R.P. Brown. 1991. Microgeographic clines in the size of mature male Gallotia galloti (Squamata: Lacertidae) on Tenerife: Causal hypotheses. Herpetologica 47:28 37. PDF

Wüster, W. & R. S. Thorpe. 1991. Asiatic cobras: systematics and snake bite. Experientia. 47:205‑209. PDF

Malhotra, A. and R. S. Thorpe. 1991. Microgeographic variation in Anolis oculatus on the island of Dominica, West Indies. J. Evol. Biol. 4:321‑335. PDF

Thorpe, R.S. 1991. Clines and cause: Microgeographic variation in the Tenerife gecko (Tarentola delalandii). Syst. Zool. 40:172‑187. PDF

Thorpe, R.S., R.P. Brown, A. Malhotra & W. Wüster, 1991. Geographic variation and population systematics: distinguishing between ecogenetics and phylogenetics. Boll. Zool.58: 329‑335. PDF

Brown, R. P., and R. S. Thorpe. 1991. Within‑island microgeographic variation in body dimensions and scalation of the skink Chalcides sexlineatus, with testing of causal hypotheses. Biol. J. Linn. Soc.  44:47‑64. PDF

Brown, R. P. and R. S. Thorpe. 1991. Within‑island microgeographic variation in the colour pattern of the skink, Chalcides sexlineatus: pattern and cause.  J. Evol. Biol. 4:557‑574. PDF

Greer, A., Thorpe, R.S. & Malhotra, A. 1991. Natural history notes from the Roe Plain, Western Australia. West Australian Naturalist. 18:178-184.


Wüster, W and Thorpe, R.S. 1990. Systematics and biogeography of the Asiatic cobra (Naja naja) species complex in the Philippine Islands. In Peters G. and R. Hutterer (Eds.) Vertebrates in the Tropics. 333‑344. Museum A Koenig, Bonn. PDF

Baez, M. and R. S. Thorpe. 1990. Analisis preliminar de las divergencias entre las poblaciones de Chalcides viridanus en las Islas Canarias. Vieraea. 19:209‑213.

Brown, R.P., R. S. Thorpe and M. Baez. 1990. Within‑island geographic variation of Chalcides in Tenerife and Gran Canaria. In Evolution and conservation on the North Atlantic Islands. Manchester.


Butler, P.M, Thorpe, R.S. & Greenwood M.  1989. Interspecific relationships of African crocidurine shrews (Mammalia:Soricidae) based on multivariate-analysis of mandibular data. Biol. J. Linn. Soc. 96:373‑412.

Wüster, W. & Thorpe, R.S. 1989. Population affinities of the Asiatic cobra (Naja naja) species complex in south-east Asia: reliability and random resampling. Biol J Linn Soc.36:391‑409. PDF

Thorpe, R.S. 1989. Geographic variation: Multivariate analysis of six character systems in snakes in relation to character number. Copeia. 1989:63‑70. PDF

Thorpe, R.S. & Brown R.P. 1989. Microgeographic variation in the colour pattern of the lizard Gallotia galloti within the island of Tenerife: distribution, pattern and hypothesis testing. Biol J Linn Soc. 38:303‑322. PDF

Corti, M. & Thorpe, R.S. 1989. Morphological clines across a karyotypic hybrid zone of house mice in central Italy. J Evol Biol. 2:253‑264. PDF

Thorpe, R.S. & Brown, R.P. 1989. Testing hypothesized causes of within‑island geographic variation in the colour of lizards. Experientia 45:397‑400. PDF

Thorpe, R.S. 1989. Pattern and function of sexual dimorphism: A biometric study of character variation in the grass snake (Natrix natrix, Colubridae) due to sex and its interaction with geography. Copeia. 1989:53‑63. PDF

Thorpe, R. S. 1989. A review of "Natural History Excursions in Tenerife. A guide to the countryside, plants and animals" by M & P Ashmole. In Journal of the British Ecological Society


Thorpe, R.S. 1988. Multiple group principal component analysis and population differentiation. J. Zool. Lond. 216:37‑40. PDF

Corti, M., Thorpe, R.S., Sola, L., Sbordoni, V. & Cataudella, S. 1988. Multivariate morphometrics in aquaculture: a case study of six stocks of the common carp (Cyprinus carpio)  from Italy. Can. J. Fish. Aquat. Sci. 45:1548‑1554. PDF


Thorpe, R.S. & Baez, M. 1987. Geographic variation within an island: univariate and multivariate contouring of scalation, size and shape of the lizard Gallotia galloti. Evolution. 41:256‑268. PDF

Thorpe, R.S. 1987. Congruence between independent character systems across a hybrid zone: patterns in geographic space. Z. Zool. Syst. Evolut- forsch. 25:161‑169. PDF

Wüster, W. & Thorpe, R.S. 1987. Geographic variation in the cobras of the genus Naja in south‑east Asia: a multivariate analysis. pp 449‑452. In J.J. Gelder, H. Strigbosch & P.J.M. Bergers, eds. Proc. 4th Ord. Gen. Meeting of the Societas Europaea  Herpetologica. PDF

Thorpe, R.S. 1987. Geographic variation: a synthesis of cause, data, pattern and congruence in relation to subspecies, multivariate analysis and phylogenesis. Boll. Zool. 54:3‑11. PDF

Searle, J.B. & Thorpe, R.S. 1987. Morphological variation in the common shrew (Sorex araneus) in relation to karyotype and geography. J. Zool. Lond. 212:373‑377.

Thorpe, R.S. 1987. Complex clines: the predictivity of complicated patterns of geographic variation portrayed by multivariate analysis. Biol. J. Linn. Soc. 31:75‑88. PDF


Thorpe, R.S. 1986. Evolution and character congruence in some western Indian Ocean Phelsuma: Numerical analysis of biochemistry, shape, and scalation. J. Zool. Lond A . 208:429‑441. PDF

 Thorpe, R. S. 1986. Population affinities and morphometry with particular reference to geographic variation in reptiles. DSc Thesis. University of Aberdeen.


Thorpe, R.S. 1985. Relative similarity between subspecies of the western Canary Island lizard, Gallotia galloti. Bonn. zool. Beitr. 36: 529‑532.

Thorpe, R.S. 1985. Extent of racial divergence in the eastern Canary island lizard, Gallotia atlantica. Bonn. zool. Beitr. 36:507‑512.

Thorpe, R.S. 1985. Alternative hypotheses for the causation of geographic variation in the western Canary Island lizard Gallotia galloti. Bonn. zool. Beitr. 36:533‑539.

Thorpe, R.S. 1985. Body size, island size and variability in the Canary Island lizards of the genus Gallotia. Bonn. zool. Beitr. 36:481‑487. PDF

Thorpe, R.S., Watt, K. & Baez, M. 1985. Some interrelationships of the Canary Island lizards of the genus Gallotia. Bonn. zool. Beitr. 36:577‑584.

Baez, M. & Thorpe, R.S. 1985. Microevolution of the lizard Gallotia galloti within the island of Tenerife. Bonn. zool. Beitr. 36:513‑515.

Thorpe, R.S. 1985. Character number and the multivariate analysis of simple patterns of geographic variation: categorical or "stepped clinal" variation. Syst. Zool. 34:127‑139. PDF

Thorpe, R.S. 1985. Clines: character number and the multivariate analysis of simple patterns of geographic variation. Biol. J. Linn. Soc. 26:201‑214. PDF

Thorpe, R.S. 1985. A numerical analysis of isoelectric focused keratin monomers: affinities of some western Indian Ocean green geckos Biochem. Syst. Ecol. 13:63‑69. PDF

Thorpe, R.S. 1985. The effect of insignificant characters on the multivariate analysis of simple patterns of geographic variation. Biol. J. Linn. Soc. 26:215‑223. PDF


Thorpe, R.S. 1984. Multivariate patterns of geographic variation between the island and mainland populations of the eastern grass snake (Natrix natrix natrix). J. Zool. Lond. 204:551‑561. PDF

Thorpe, R.S. 1984. Primary and secondary transition zones in speciation and population differentiation: a phylogenetic analysis of range expansion. Evolution. 38:233‑243. PDF

Thorpe, R.S. 1984. Coding morphometric characters for constructing Distance Wagner networks. Evolution. 38:244‑255. PDF

Leamy, L. & Thorpe, R.S. 1984. Morphometric studies in inbred and hybrid house mice. Heterosis, homeostasis and heritability of size and shape. Biol. J. Linn. Soc. 22:233‑241. PDF

Thorpe, R.S. 1984. Geographic variation in the western grass snake (Natrix natrix helvetica) in relation to hypothesized phylogeny and conventional subspecies. J.Zool. Lond. 203:345‑355. PDF

Boyle, P.R. & Thorpe, R.S. 1984. Optic gland enlargement and female gonad maturation in a population of the octopus Eledone cirrhosa: a multivariate analysis. Marine Biol. 79:127‑132. PDF


Thorpe, R.S. & Giddings, M.R. 1983. The biochemistry of keratin in green geckos (Phelsuma) and its systematic value. Herpetologica. 39:399‑410. PDF

Thorpe, R.S. 1983. A biometric study of the effects of growth on the analysis of geographic variation: tooth number in green geckos (Reptilia: Phelsuma). J. Zool. Lond. 201:13‑26. PDF

Thorpe, R.S. & Leamy, L. 1983. Morphometric studies in inbred and hybrid house mice (Mus sp): a multivariate analysis of size and shape. J. Zool. Lond. 199:421‑432. PDF

Thorpe, R.S. 1983. Phylogenetic analysis of range expansion in the grass snake: reticulate evolution: primary and secondary contact zones. Numerical Taxonomy: Proceedings of a NATO Advanced Study Institute. NATO Advanced Study Institute Series G (Ecological Sciences), No.1, ed. Felsenstein, J. , Berlin, Heidelberg and New York (Springer‑Verlag) pp. 464‑468.

Thorpe, R.S. 1983. A review of the numerical methods for recognising and analysing racial differentiation. Numerical Taxonomy: Proceedings of a NATO Advanced Study Institute. NATO Advanced Study Institute Series G (Ecological Sciences), No.1, ed. Felsenstein, J., Berlin, Heidelberg and New York (Springer‑Verlag) pp. 404‑423. PDF

Thorpe, R.S. 1983. Research into the evolution and taxonomy of snakes and lizards. Bull. Brit. Herp. Soc. 7:30‑31.


Thorpe, R.S. 1982. Reticulate evolution and cladism: tests for the direction of evolution. Experientia. 38:1242‑1243. PDF

Thorpe, R.S. 1982. Molossid bat morphometry. A review of "A multivariate study of the family Molossidae (Mammalia, Chiroptera): morphology, ecology, evolution". Freeman, P.U. Fieldiana Zoology, New Series No.7 Field Museum of Natural History, Chicago, 173 pp.1981. J. Mamm. 63:353‑354.

Thorpe, R.S., Corti, M. & Capanna, E. 1982. Morphometric divergence of Robertsonian populations/species of Mus: a multivariate study of size and shape. Experientia. 38:920‑923. PDF


Thorpe, R.S. 1981. Racial divergence and subspecific status of the Gotland grass snake: a comment on Nilson and Andren's paper. Zool. J. Linn. Soc. 72:369‑370.

Thorpe, R.S. & Giddings, M.R. 1981. A novel biochemical systematic technique for herpetology based on epidermal keratin. Experientia, 37:700‑702. PDF

Thorpe, R.S. 1981. The morphometrics of the mouse: A review. In The Biology of the House Mouse, Ed. R.J. Berry. Symp. Zool. Soc. Lond., 47: 85‑125.


Crawford, CM. & Thorpe, R.S. 1980. Identification, morphometrics an cytogenetics of green geckos (Phelsuma) from Praslin, Seychelles (with notes on P. longinsulae, Mahe). Biol. J. Linn. Soc. 15:29‑37.

Thorpe, R.S. 1980. A comparative study of ordination techniques in numerical taxonomy in relation to racial variation in the ringed snake Natrix natrix (L.). Biol. J. Linn. Soc. 13:7‑40. PDF

Thorpe, R.S. 1980. Microevolution and taxonomy of European reptiles with particular reference to the grass snake Natrix  natrix and the wall lizards Podarcis sicula and P.  melisellensis. Biol. J. Linn. Soc. 14:215‑233. PDF

Thorpe, R.S. 1980. Racial affinities and geographic variation: comparison of the use of multivariate morpometrics and conventional subspecies. Proc. Euro. Herp. Symp. Oxford, 5‑6.

Miller, P., Tawil, E., Thorpe, R.S. and Webb, C. 1980. Haemoglobins and the systematic problems set by Gobioid fishes. pp 195‑233 In Chemosystematics: Principles and Practice. Eds. Bisby, Vaughan and Wright. Academic Press, London.


Crawford, CM. & Thorpe, R.S. 1979. Body temperatures of two geckos (Phelsuma) and a skink (Mabuya) in Praslin, Seychelles. Brit. J. Herpetol. 6:25‑31.

Thorpe, R.S. & Crawford, CM. 1979. The comparative abundance and resource partitioning of two green geckos species (Phelsuma) on Praslin, Seychelles. Brit. J. Herpetol. 6:19‑24.

Thorpe, R.S. 1979. Multivariate analysis of the population systematics of the ringed snake, Natrix natrix (L). Proc. Royal Soc. Edin. 78B:1‑62. PDF


Thorpe, R.S. & McCarthy, C.J. 1978. A preliminary study, using multivariate analysis, of a species complex of African house snakes (Boaedon fuliginosus). J. Zool. Lond. 184:489‑506.


Thorpe, R.S. 1976. Biometric analysis of geographic variation and racial affinities. Biological Reviews. 51:407‑452. PDF


Thorpe, R.S. 1975. Biometric analysis of incipient speciation in the ringed snake,  Natrix natrix (L.). Experientia. 31:180‑182. PDF

Thorpe, R.S. 1975. Quantitative handling of characters useful in snake systematics with particular reference to intraspecific variation in the ringed snake Natrix natrix (L.). Biol. J. Linn. Soc. 7:27‑43. PDF


Thorpe, R.S. 1973. Intraspecific variation of the ringed snake Natrix natrix (L.). Ph.D. Thesis.